For a full list of publications, visit eSpace


Dscam2 suppresses synaptic strength through a PI3K-dependent endosomal pathway.
Odierna GL, Kerwin SK, Harris LE, Shin GJ, Lavidis NA, Noakes PG, Millard SS.
J Cell Biol. 2020 Jun 1;219(6):e201909143. doi: 10.1083/jcb.201909143.
PMID: 32259198

Elevated levels of Drosophila Wdr62 promote glial cell growth and proliferation through AURKA signalling to AKT and MYC.
Shohayeb B, Mitchell N, Millard SS, Quinn LM, Ng DCH.
Biochim Biophys Acta Mol Cell Res. 2020 Jul;1867(7):118713. doi: 10.1016/j.bbamcr.2020.118713. Epub 2020 Apr 1.
PMID: 32246948

The association of microcephaly protein WDR62 with CPAP/IFT88 is required for cilia formation and neocortical development.
Shohayeb B, Ho U, Yeap YY, Parton RG, Millard SS, Xu Z, Piper M, Ng DCH.
Hum Mol Genet. 2020 Jan 15;29(2):248-263. doi: 10.1093/hmg/ddz281.
PMID: 31816041


Li, JS and Millard SS (2019). Deterministic splicing of Dscam2 is regulated by Muscleblind. Sci Adv 5(1): eaav1678.


Millard SS, Pecot MY (2018). Strategies for assembling columns and layers in the Drosophila visual system. Neural Development 13(1):11.

Kerwin SK, Li JS, Noakes PG, Shin GJ, Millard SS (2018). Regulated alternative splicing of Drosophila Dscam2 is necessary for attaining the appropriate number of photoreceptor synapses. Genetics 208 (2): 717-28.


Lim NR, Shohayeb B, Zaytseva O, Mitchell N, Millard SS, Ng DCH, and Quinn LM (2017). Glial-specific functions of microcephaly protein WDR62 and interaction with the mitotic kinase AURKA are essential for Drosophila brain growth. Stem Cell Reports 9: 1-10.


Tadros W, Xu S, Akin O, Yi CH, Shin GJ, Millard SS and Zipursky SL (2016). Dscam proteins direct dendritic targeting through adhesion. Neuron 89 (3):480-93.


Li JS, Shin GJ, Millard SS (2015). Neuronal cell-type-specific alternative splicing: A mechanism for specifying connections in the brain? Neurogenesis 2:1, e1122699: 1-5.

Bosch DS, van Swinderen B and Millard SS (2015). Dscam2 affects visual perception in Drosophila melanogaster. Front. Behav. Neurosci. 9:149. doi: 10.3389/fnbeh.2015.00149.


Lah GJ, Li JS, Millard SS (2014).  Cell-specific alternative splicing of Drosophila Dscam2 is crucial for proper neuronal wiring.  Neuron 17;83(6):1376-88.


Li Q, Ha TS, Okuwa S, Wang Y, Wang Q Millard SS, Smith DP and Volkan-Cayirlioglu P (2013). Combinatorial rules of precursor specification underlying olfactory neuron diversity. Current Biol. 23, 2481-2490.

Paulk A, Millard SS, van Swinderen B.  (2013).  Vision in Drosophila:  Seeing the World Through a Model's Eyes.  Ann. Rev. Entomol. 58:313-32. 


Millard SS, Lu Z, Zipursky SL, Meinertzhagen IA. (2010) Drosophila Dscam proteins regulate postsynaptic specificity at multiple-contact synapses. Neuron 9;67(5):761-8.


Millard SS, Zipursky SL. (2008) Dscam-mediated Repulsion Controls Tiling and Self-avoidance. Current Opinion in Neurobiology 18(1):84-9.

Hattori D, Millard SS, Wojtowicz W, Zipursky SL. (2008) Dscam-mediated Cell Recognition Regulates Neural Circuit Formation. Annual Reviews in Cell and Developmental Biology 24:597-620.


Millard SS, Flanagan JJ, Pappu KS, Wu W, Zipursky SL.  (2007) Dscam2 mediates axonal tiling in the Drosophila visual system.  Nature 447(7145):720-4.